A compound attribute that is the ratio of amount of guanylin (human) in blood to amount of galectin-4 (human) in blood. The trait is GUCA2A/LGALS4 protein level ratio in blood (OBA_2053456, an Ontology of Biological Attributes entry). Also known as: the ratio of amount of guanylin (human) in blood to amount of galectin-4 (human) in blood.