A compound attribute that is the ratio of amount of proline-rich AKT1 substrate 1 (human) in blood to amount of forkhead box protein O1 (human) in blood. The trait is AKT1S1/FOXO1 protein level ratio in blood (OBA_2054286, an Ontology of Biological Attributes identifier). Also known as: the ratio of amount of proline-rich AKT1 substrate 1 (human) in blood to amount of forkhead box protein O1 (human) in blood.