A compound attribute that is the ratio of amount of carbonic anhydrase 4 (human) in blood to amount of tumor necrosis factor ligand superfamily member 12 (human) in blood. The trait is OBA_2054821 (CA4/TNFSF12 protein level ratio in blood). Also known as: the ratio of amount of carbonic anhydrase 4 (human) in blood to amount of tumor necrosis factor ligand superfamily member 12 (human) in blood.