As part of the intermicrovillar adhesion complex/IMAC plays a role in epithelial brush border differentiation, controlling microvilli organization and length. Acts as a light chain for MYO7B and is required for efficient targeting of the IMAC to the tips of border brush microvilli. . This is the function of CALML4 (calmodulin like 4, Ensembl gene identifier ENSG00000129007).