Major component of the transverse central element of synaptonemal complexes (SCS), formed between homologous chromosomes during meiotic prophase. Requires SYCP1 in order to be incorporated into the central element. May have a role in the synaptonemal complex assembly, stabilization and recombination (By similarity). . This is the function of SYCE2 (synaptonemal complex central element protein 2, Ensembl gene identifier ENSG00000161860).