Involved in early stages DNA double-strand break (DSB) repair via homologous recombination (HR) (PubMed:16710300, PubMed:21965664, PubMed:34253720, PubMed:39169038, PubMed:40345587). Forms a heterodimeric complex with ZSWIM7/SWS1 that promotes HR by regulating replication protein-A (RPA) dynamics on single-stranded DNA, thereby stabilizing RAD51 and DMC1 filaments on DNA (PubMed:34253720, PubMed:39169038, PubMed:40345587). The SWSAP1-ZSWIM7/SWS1 heterodimer is essential during meiosis by stabilizing DMC1 filaments (PubMed:32719396, PubMed:33713115, PubMed:34402903). In contrast, it is critical for only certain types of homologous-directed DNA repair, such as inter-homolog homologous recombination (IR-HR) (PubMed:34253720). IR-HR is a DNA repair mechanism where a cell repairs a DSB on one chromosome using the matching homologous chromosome as a template, rather than the sister chromatid (PubMed:34253720). {ECO:0000269|PubMed:16710300, ECO:0000269|PubMed:21965664, ECO:0000269|PubMed:32719396, ECO:0000269|PubMed:33713115, ECO:0000269|PubMed:34253720, ECO:0000269|PubMed:34402903, ECO:0000269|PubMed:39169038, ECO:0000269|PubMed:40345587}. This is the function of Ensembl gene identifier ENSG00000214941 (ZSWIM7, zinc finger SWIM-type containing 7).